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Cancer pagurus, commonly known as the edible crab or brown crab, is a species of found in the , North , and perhaps the Mediterranean Sea. It is a robust crab of a reddish-brown colour, having an oval with a characteristic "pie crust" edge and black tips to the claws. A mature adult may have a carapace width up to and weigh up to . C. pagurus is a nocturnal predator, targeting a range of molluscs and crustaceans. It is the subject of the largest in , centred on the coasts of the Ireland and Britain, with more than 60,000 tonnes caught annually.


Description
The of C. pagurus adults is a reddish-brown colour, while in young specimens it is purple-brown. It occasionally bears white patches, and is shaped along the front edge into nine rounded lobes, resembling a pie crust. Males typically have a carapace long, and females long, although they may reach up to long in exceptional cases. Carapace width is typically , or exceptionally up to . A fold of the carapace extends ventrally to constitute a branchial chamber where the lie.
(1997). 9780412710605, Cambridge University Press.

The first pereiopod is modified into a strong cheliped (claw-bearing leg); the claw's fingers, the and propodus, are black at the tips. The other pereiopods are covered with rows of short stiff ; the dactylus of each is black towards the tip, and ends in a sharp point.

From the front, the antennae and are visible. Beside these, the orbits are where the are situated. The mouthparts comprise three pairs of , behind which are a pair of maxillae, a pair of maxillules, and finally the mandibles.


Life cycle
Reproduction occurs in winter; the male stands over the female and forms a cage with his legs protecting her while she . Internal fertilisation takes place before the hardening of the new carapace, with the aid of two abdominal appendages (gonopods). After mating, the female retreats to a pit on the sea floor to lay her eggs. Between 250,000 and 3,000,000 fertilised eggs are held under the female's abdomen up to eight months until they hatch.

The first developmental stage after hatching is a larva (1 mm) called the that develops into a (), and finally a juvenile. The first juvenile stage is characterised by a well-developed abdomen, which in time becomes reduced in size and folded under the sternum. Juveniles settle to the sea floor in the , where they stay until they reach a width of , and then migrate to deeper water. The growth rate in males slows from an increase in carapace width of 10 mm per year before it is 8 years old, to 2 mm per year thereafter. Females grow at about half the rate of males, probably due to the energetic demands of egg laying. is reached at a carapace width of in females, and in males. is typically 25–30 years, although exceptional individuals may live up to 100 years.


Distribution and ecology
C. pagurus is abundant throughout the northeast Atlantic as far as in the north and in the south, on mixed coarse grounds, mud, and sand from the shallow to depths around . It is frequently found inhabiting cracks and holes in rocks, but occasionally also in open areas. Smaller specimens may be found under rocks in the . Unconfirmed reports suggest that C. pagurus may also occur in the Mediterranean Sea and .

Adults of C. pagurus are , hiding buried in the substrate during the day, but foraging at night up to from their hideouts. Their diet includes a variety of (including the crabs and Pilumnus hirtellus, the Porcellana platycheles and Pisidia longicornis, and the Galathea squamifera) and (including the and Littorina littorea, and the , , Cerastoderma edule, , and Lutraria lutraria). It may stalk or ambush motile prey, and may dig large pits to reach buried molluscs. The main of C. pagurus is the , which even attacks them inside the crab pots that fishermen use to trap them.

(2025). 9780761472667, Marshall Cavendish.


Diseases
Compared to other commercially important crab species, relatively little is known about diseases of C. pagurus. Its parasites include , such as the white spot syndrome virus, various that cause dark on the , and -like that cause "pink crab disease". Other microscopic include , , , and . C. pagurus is also targeted by parasites, including and parasitic . A number of sessile animals occasionally settle as on the exoskeleton of C. pagurus, including barnacles, , such as Janua pagenstecheri, , and saddle oysters.


Fishery
C. pagurus is heavily throughout its range, being the most commercially important crab species in Western Europe. The crabs are caught using crab pots (similar to ), also known as creels, which are placed offshore and baited. The catch of C. pagurus has increased steadily, rising from 26,000 tonnes in 1978 to 60,000 t in 2007, of which more than 70% was caught around the . The fishery is widely dispersed around the British and Irish coasts, and C. pagurus is thought to be across much of this area.
(2009). 9781906634360, Sea Fish Industry Authority. .
Most of the edible crabs caught by the British fleet are exported live for sale in and .

A number of legal restrictions apply to the catching of C. pagurus. Catching "berried" crabs (females carrying eggs) is illegal, but since ovigerous females remain in pits dug in the sediment and do not feed, fishing pressure does not affect the supply of larvae. Minimum landing sizes (MLSs) for C. pagurus are set by both the technical regulations and by the UK government. Different minimum sizes are employed in different geographical areas, to reflect differences in the crab's growth rate across its range. In particular, the " crab" fishery along the coasts of , and is subject to an MLS of , rather than the MLS in most of the species' range. An intermediate value of is used in the rest of the North Sea between the 56°N and the border, and in the south of 55°N. Around , , and the Isles of Scilly, the MLS for males is different () from females (). The Norwegian catch is 8,500 tons annually, compared to 20,000 tons in the United Kingdom, 13,000 tons in Ireland, 8,500 tons in France, and a total 45,000 tons globally. Recent studies have shown that edible crabs are negatively affected by electromagnetic fields emitted from sub-sea power cables around offshore wind farms.


Cookery
Around one-third of the weight of an adult edible crab is meat, of which one-third is white meat from the claws (see declawing of crabs), and two-thirds is white and brown meat from the body.
(2025). 9780192806819, Oxford University Press.
As food, male edible crabs are referred to as cocks and females as hens. Cocks have more sweet white meat; hens have more rich brown meat. Dishes include (crab meat arranged in the cleaned shell, sometimes with decoration of other foodstuffs), soups such as bisque or , pâtés, , and hot soufflés.
(1996). 9780748725663, .


Taxonomy and systematics
According to the rules of the International Code of Zoological Nomenclature, Cancer pagurus was first described by in 1758, in the tenth edition of his , which marks the starting point of zoological nomenclature. It was chosen to be the of the genus Cancer by Pierre André Latreille in 1810. The specific epithet is a word, deriving from the (), which, alongside (), was used to refer to edible marine crabs; neither classical term can be confidently assigned to a particular species.
(2025). 9780415232593, .

Although the genus Cancer formerly included most crabs,

(2025). 9780643069060, .
it has since been restricted to eight species. Within that set of closely related species, the closest relative of C. pagurus is the , C. borealis, from the east coast of .


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